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The Northwest Coast of North America stretches 4000 km from Bering Strait to Washington State. Here we review the history of glaciation, sea level, oceanography, and climate along the Northwest Coast and in the subarctic Pacific Ocean during the Last Glacial Maximum and deglaciation. The period of interest is Marine Isotope Stage 2 between ca. 29,000 calendar years ago (29 ka) and 11,700 calendar years ago (11.7 ka). The glacial history of the Northwest Coast involved multiple glacial systems responding independently to latitudinal variations in climate caused by changes in the North American ice sheets and in the tropical ocean-atmosphere system. Glaciers reached their maximum extents 1–5 kyrs later along the Northwest Coast than did large sectors of the Laurentide and Fennoscandian Ice Sheets. Local, Last Glacial Maxima were reached in a time-transgressive, north to south sequence between southwestern Alaska and Puget Sound. The history of relative sea level along the Northwest Coast during Marine Isotope Stage 2 was complex because of rapid isostatic adjustments by a thin lithosphere to these time-transgressive glacial fluctuations. Multiple lines of evidence suggest Bering Strait was first flooded by the sea after 11 ka and that it probably did not assume its present-day oceanographic functions until after 9 ka. The coldest intervals occurred during Heinrich Event 2 (ca. 26–23.5 ka), again between ca. 23 and 21.5 ka, and during Heinrich Event 1 (ca. 18–15 ka). During these times, mean annual sea surface temperatures cooled by 5o to 8o C in the Gulf of Alaska, and glacial equilibrium-line altitudes fell below present sea level in southern Alaska and along the Aleutian Island chain. Sea ice episodically expanded across the subarctic Pacific in winter. Oceanographic changes in the Gulf of Alaska tracked variations in the vigor of the Asian Summer Monsoon. The deglaciation of the Northwest Coast may have served as the trigger for global climate changes during deglaciation. Starting ca. 21 ka, marine-based glaciers there were increasingly destabilized by rising eustatic sea level and influxes of freshwater and heat associated with the rejuvenation of the Asian Summer Monsoon. Rapid retreat of marine-based glaciers began ca. 19 ka and released large numbers of ice bergs and vast amounts of freshwater into the Northeast Pacific. Resultant cooling of the North Pacific may have been teleconnected to the North Atlantic through the atmosphere, where it slowed Atlantic Meridional Overturning Circulation and initiated the global effects of Heinrich Event 1, ca. 18–15 ka. During the Younger Dryas, ca. 12.8–11.7 ka, mean annual sea surface temperatures were 4o to 6o C cooler than today in the Gulf of Alaska, and sea ice again expanded across the subarctic Pacific in winter. Conditions of extreme seasonality characterized by cold, dry winters and warm, steadily ameliorating summers caused by the southward diversion of the Aleutian Low in winter may explain the previously enigmatic records of Younger Dryas climate along the Northwest Coast. 

Climate warming can cause arthropods to express plastic and/or evolved changes in morphology. Previous studies have demonstrated that body sizes of Arctic butterflies are influenced by the temperatures experienced as larvae. To investigate whether this was occurring among Alaskan butterflies, we analyzed temporal trends in the wing sizes of three Holarctic species,Colias hecla,Boloria charicleaandBoloria freija, using museum specimens collected in Arctic tundra regions of Alaska between 1971 and 1995. Wing length was compared to accumulated growing degree days (GDD) during both the spring of the year collected and the previous year's summer during the normal period of larval development. We used mixed‐effects models to test if spring and summer temperatures affected adult morphology. Results show that for every 1°C increase in average seasonal temperature, wingspans decreased between 0.7 and 5 mm, withB. freijathe most strongly affected. Our results suggest that the morphological sensitivity of Arctic butterflies to warming is the outcome of interactions between life‐history traits and regional climate, with all species sensitive to warming the summer before the flight year as well as warming the spring of the flight year.Boloria freija, which overwinters as late instar larvae that do not feed before pupation the following spring, was particularly strongly affected by summer warming. 

Abstract Winter is a critical season for land‐surface feedbacks and ecosystem processes; however, most high‐latitude paleo‐environmental reconstructions are blind to cold season conditions. Here we introduce a winter‐sensitive, paleo‐proxy record that is based on the relative frequency of tangential rows of traumatic resin ducts (TRDs) in the annual growth rings of mountain hemlocks (Tsuga mertensiana) growing near treeline in Southeast Alaska. Hemlocks produce a row of TRDs in the earlywood portion of their annual rings in response to cambial damage incurred during winter. Multidecadal bouts of TRD production were followed by growth‐leader replacement, reaction wood formation, and divergence in radial growth between storm‐damaged trees and less exposed mountain hemlock forests. These patterns are consistent with TRDs being a response to tree damage caused by ice and snowstorms, a conclusion supported by the krummholz tree architecture at these sites. This relationship is further corroborated by significant correlations between our TRD record and the strength of the wintertime Aleutian Low (AL) pressure system that is linked to tree‐damaging agents like wind, precipitation, and ice storm strength in Southeast Alaska. The combined TRD/krummholz architecture record indicates that abrupt shifts between strong and weak AL phases occurred every several decades since CE 1700 and that the 1800s had relatively long AL phases with heavy snowpacks. In addition to describing the magnitude and tempo of wintertime climate change in Northwestern North America, these results suggest that North Pacific Decadal Variability underlies the long‐term dynamics of treeline ecosystems along the northeast Pacific coast.